Biology (BIOL) Courses
College of Arts and Sciences
BIOL 101 General Biology (3) A one-semester introductory biology course for non-majors.
BIOL 101L Gen Biol Lab (1) (lab) Laboratory for General Biology. (Optional but recommended)
BIOL 125 Intro Cell & Molecular Biol (3) Introduction to cell and molecular biology for majors in the natural and health sciences. Cell structure and function, the molecular basis of life, metabolic pathways, classical and molecular genetics, the molecular and biological evolution of the universe. Pre: high school biology (or BIOL 101), high school chemistry (or CHEM 114).
BIOL 156 Nat Hist & Conservatn Hawn Isl (3) The formation of the Hawaiian Islands, establishment of their native terrestrial and marine flora and fauna, and human impacts and conservation. (Same as MARE 156). (GenEd/IntReq: H/A/P, HPP)
BIOL 156L Nat History Field Trips (1) (lab) Field trips for Natural History and Conservation of the Hawaiian Islands. (Same as MARE 156L). (GenEd/IntReq: H/A/P)
BIOL 171 Marine Biology-Diversity (3) Marine organisms: classification, structure, physiology, ecology and adaptations to the marine environment. This course satisfies CAS general education requirements in the Natural Sciences. (Same as MARE 171.)
BIOL 171L Marine Biology Laboratory (1) (lab) Provides students with direct exposure to the biota of Hawaiʻi via laboratory work and field trips to sites around Hilo. The course focuses on the identification, natural history, and ecology of common marine organisms. Pre: current or previous enrollment in BIOL/MARE 171. (Same as MARE 171L).
BIOL 175 Introductory Biology I (3) Principles of cell structure, replication, and metabolism. Classical and molecular genetics, and evolution. Biodiversity of prokaryotes, viruses, fungi, and plants. Plant structure and function. Biology 175 and 176 are offered both semesters, and students may enroll in either (but not both) during the fall or spring semester.
BIOL 175L Introductory Biology I Lab (1) (lab) Laboratory for Introductory Biology I. Laboratory exercises covering cell structure, replication, and metabolism; classical and molecular genetics; evolution; and biodiversity of prokaryotes, viruses, fungi, and plants. BIOL 175L should be taken concurrently with BIOL 175.
BIOL 176 Introductory Biology II (3) Biodiversity of animal-like protistans, invertebrates, and vertebrates. Animal tissues, sensory reception and integration, endocrine systems, support and movement, circulation and immunity, gas exchange, digestion, kidney function, reproduction and development. Population and community ecology, energy flow and biogeochemical cycles. BIOL 175 and 176 are each taught both semesters, and students may enroll in either (but not both) during either fall or spring semester.
BIOL 176L Introductory Biology II Lab (1) (lab) Laboratory for Introductory Biology II. Laboratory exercises covering structure, function, and natural history of animal-like protistans, invertebrates and vertebrates; structure and function of animal tissues; reproduction and development; and community ecology. BIOL 176L should be taken concurrently with BIOL 176.
BIOL 190 Hawn Marine Field Experience (2) Provides a unique opportunity for students to experience the marine environment of the Big Island. Course consists of field trips to coastal and underwater sites around the island. Some swimming and snorkeling involved but not required. (Same as MARE 190). (GenEd/IntReq: H/A/P)
BIOL 205 Gen Entomology (3) Structure, classification and identification of insects. Pre: BIOL 175 or 176 or instructor’s consent. (Same as ENTO 304).
BIOL 254 Fund Animal Nutrition (3) Comparative animal digestive systems and metabolism. Essential nutrients, their functions, mechanisms of action and interrelationships. Pre: ANSC 141, CHEM 124 and 125, or instructor’s consent. (Same as ANSC 254).
BIOL 275 Fund Microbiology (3) A survey or microbiology with emphasis on bacteria, viruses, infectious diseases and their control. Pre: one semester of college chemistry. Recommended: at least one semester of BIOL 101, 175 or 176. (GenEd/IntReq: GCC)
BIOL 275L Microbiology Lab (1) (lab) Required laboratory for Fundamentals of Microbiology.
BIOL 280 Biostatistics (3) Statistical analysis as applied to research in the Biological Sciences. Theory and applications of statistics; experimental design; basic statistical concepts; hypothesis testing; parametric and non-parametric analyses. Group and independent projects, computer analysis of data.
BIOL 309 Biogeography (3) Basic evolutionary and ecological principles underlying the dynamics of plant and animal populations. Mechanisms of isolation, speciation, dispersal, migration, and competition as they affect past and present world distribution patterns. Island biogeography. Pre: GEOG 101; BIOL 101 or 175 or 176; or instructor's consent. (Same as GEOG 309). (GenEd/IntReq: H/A/P)
BIOL 323 Mammal Physiology (3) Structure and function of the animal body. A general study of anatomy, but emphasis placed on understanding the physiology. (Same as ANSC 350)
BIOL 357 Evolution (3) Organic evolution as a unifying theory of biology. Topics include the history of ideas of evolution, adaptation of populations, genetic drift, molecular evolution and the neutral theory, quantitative genetics, speciation and phylogeny, biogeography and macro-evolutionary trends, and mass extinctions. Pre: BIOL 125 or 175 or 176.
BIOL 357L Evolutionary Genetics Lab (1) (lab) A laboratory and field course to be taken in conjunction with BIOL 357 lecture. Laboratory exercises will introduce students to the techniques in evolutionary genetics with special reference to Hawaiian plants and animals. Topics to be covered include: quantitative genetics, natural selection molecular genetics analysis of populations and species, ecological genetics and adaptation, and conservation genetics. Pre: BIOL 270, BIOL 270L and BIOL 280.
BIOL 360 Marine Resources (3) A survey of human use of the marine environment including physical and biological resources. Topics covered include: fisheries, mariculture, marine mineral and energy resources, chemical resources of sea water, the use of coastal lands and waste disposal in the sea. Pre: MARE 201 or BIOL/MARE 171, or instructor’s consent. (Same as MARE 360).
BIOL 366 Trop Marine Rsrch Investigatn (3) Research projects on marine-related problems. Students will do a literature search; develop experimental design; collect, reduce and analyze data; do a written final report; and present findings at a symposium. Projects will be selected from a list of topics or can be original with the consent of the instructor. Pre: instructor’s consent. (Same as MARE 366).
BIOL 371 Biology Of Marine Invertebrate (3) A survey of the major groups of invertebrates focusing on those dominant in the marine environment. Students will learn methods used to identify and classify invertebrates and will survey the anatomy, physiology, and natural history of the major groups. Pre: MARE 265 or BIOL 176 or their equivalent, concurrent enrollment in BIOL/MARE 371L. (Same as MARE 371).
BIOL 371L Bio Of Marine Invertebrate Lab (1) (lab) Direct exposure to the major groups of invertebrates in marine environment, focusing on those present in Hawaiʻi. Students will learn to identify and classify invertebrates and will survey the anatomy and natural history of the major groups. Pre: BIOL/MARE 371 or concurrent enrollment. (Same as MARE 371L).
BIOL 375 Biology of Microorganisms (3) Fundamental principles of microbiology combined with recent developments in and applications to microbial evolution, ecology, molecular genetics and immunology. Pre: BIOL 270 and 270L.
BIOL 375L Biology of Microorganisms Lab (1) (lab) Required laboratory for BIOL 375 covering fundamental principles of microbiology (culturing and identification). Supplemental components include specific labs in microbial ecology, molecular immunology, and genetics. Pre: BIOL 270 and 270L.
BIOL 392 Biology & Philosophy (3) Philosophical examination of the implications of modern biology for how we understand ourselves and our relations to the natural world. Evolutionary, genetic, developmental, and ecological topics will be discussed. Pre: previous work in Philosophy or Biology, or instructor’s consent. (Same as PHIL 392).
BIOL 415L Cell Biology Lab (2) (lab) Light and electron microscopy of selected cells. Optional laboratory for Cell Biology. Pre: concurrent enrollment in BIOL 415 and completion of BIOL 270L or instructor’s consent.
BIOL 417 Plant Anatomy (4) Plant structure in relation to cultural practices, functions genetic factors and development. Pre: BIOL 175. (Same as HORT 437).
BIOL 425 Water Qual & Aquatic Product (3) Study of water quality and aquatic productivity as it relates to aquaculture and fisheries. Pre: CHEM 124 or instructor’s consent. (Same as AQUA 425).
BIOL 436 Animal Cognition (3) A survey of the historical and contemporary scientific literature on animal cognition using a wide variety of species. The course covers a broad array of topics that may include concept formation, memory processes, numerical competence, social learning and imitation, self-awareness, theory of mind, referential communication and grammatical skills. Pre: PSY 213, PSY 214, and PSY 314 or 350 or instructor's consent. (Same as PSY 436)
BIOL 437 Marine Mammal Behavior (3) An introduction to marine mammals with emphasis on the behavior of marine mammals. Special attention given to those species found in Hawaiian waters. Individual species are examined within a comparative framework. Topics include: behavioral ecology, social behavior, cetacean societies, mating systems, communication, sensory perception, and cognition. Pre: PSY 213, PSY 214, and PSY 314 or PSY 435 or instructor's consent. (Same as PSY 437)
BIOL 443 Ecological Animal Physiology (3) Study of the physiological adaptations of animals to environmental variation. The focus will be on how physiological responses to environmental factors determine the geographic distribution of animals. The course will cover the physiological and biochemical mechanisms that permit animals to adapt to potentially stressful environmental conditions, and topics will include food acquisition and digestion, energy allocation, thermal energetics, respiratory gas exchange, activity metabolism, and osmoregulation. Pre: course in basic cell biology: BIOL 125, BIOL 270, or MARE 172; or instructor’s consent.
BIOL 445 Behavioral Ecology & Evolution (3) Principles of behavioral ecology and evolution with a focus on conservation biology. Research techniques in behavioral ecology related to analyzing populations in geographically and age-structured populations. The importance of reproductive strategies, habitat selection, foraging behavior, parental care, social organizations, and the importance of migration and movement patterns on the regulation of population sizes and evolution. Population and quantitative genetics as it relates to evolution, speciation, and biodiversity. Pre: BIOL 281 and BIOL 357 or permission of instructor.
BIOL 450 Physiology Reproduction (3) Livestock reproductive anatomy and physiology. Pre: ANSC 141. Recommended: ANSC 350. (Same as ANSC 450).
BIOL 455 Plant Ecology (3) Interactions between plants and the environment at the physiological, population, community, ecosystem scales. Discussion of factors affecting plant distribution and abundance from local to global scales. Plant and people interactions will be noted in the context of our changing planet. Key theories, quantitative measurements, experimentation, and critical analysis will be emphasized through a discussion of primary scientific literature. Pre: BIOL 281 or permission of instructor.
BIOL 457 Vegetation of the Hawaiian Isl (3) This course explores the major types of vegetation in Hawaiʻi with respect to physical environments and important species. Basic concepts in plant taxonomy and identification are emphasized. Coursework includes field excursions at various sites around the island of Hawaiʻi to develop methods in surveying and monitoring vegetation. This course also examines major human impacts and conservation challenges. Pre: BIOL 156 or BIOL 281. (Same as ENSC 457).
BIOL 460 Plant Diversity & Evolution (3) The evolution of land plants from mosses to flowering plants in the context of broad-scale environmental changes since the Silurian Period. Origins and diversification of the major lineages of extant and fossil land plants, including transitions in morphology, physiology, and life-cycles associated with adaptive radiations. Hawaii's spectacular plant radiations, and molecular and phylogenetic methods used in reconstructing the evolutionary history of land plants emphasized. Pre: BIOL 357 or permission of instructor.
BIOL 467 Ecological Genetics (3) The class focuses on the genetics of whole populations and species. We will examine barriers to gene flow that influence the population structure and distribution of species. The goal of the class is to investigate how genetics informs understanding of the evolution of ecosystems and the dynamics of genetic interactions.
BIOL 481L Ecology & Evolutn Resrch Methd (2) (lab) Intensive field-laboratory supporting BIOL 481. Research topics selected from current fields of active research within ecology and evolution. Develop research hypotheses, gather data from field sites, analyze and interpret data and write reports in the style of scientific papers. Pre: concurrent enrollment in BIOL 481 required.
BIOL 482H Honors Appl Of Ecol & Evol (3) Practical experience in performing research projects in Ecology, Evolution and Conservation Biology. Students will submit project proposals for evaluation and approval, do a thorough literature review, develop an experimental design, and collect and analyze data. Students will also prepare a final written report and give a 15-minute seminar presentation on their projects. Pre: BIOL 443L, BIOL 481 and 481L; 3.5 GPA or instructor’s consent with departmental approval.
BIOL 484 Biology Of Fishes (3) The biology of marine and freshwater fishes. Topics covered include: general anatomy, locomotion, respiration, osmoregulation, sensory systems, reproduction, electrosensitive and electrogenic fishes, coloration and bioluminescence in fishes, genetic interrelationships. Pre: C- or better in BIOL/MARE 171 or BIOL 176 or their equivalent; C- or better in MARE 265 or equivalent; or instructor’s consent. (Same as MARE 484)
BIOL 484L Biology Of Fishes Laboratory (1) (lab) Anatomy of jawless, cartilaginous and bony fishes. Review of common local reef fishes. Optional laboratory and field trips for Biology of Fishes. (Same as MARE 484L).
BIOL 495A Biology Seminar (1) Lectures, discussions and research reports of topics in biology presented by faculty, students, and visiting scholars. Students attend seminars and receive CR/NC grade for the course.
BIOL 495B Biology Seminar (1) Lectures, discussions, and research reports of topics in biology presented by faculty, students and visiting scholars. Each student must attend seminars and present a talk to receive a CR/NC grade for the course.
BIOL 496 Tchg Asstance & Tutorg in Biol (1-3) (lab) Practice in individual tutoring, and in the preparation and presentation of selected topics in Biology lecture or laboratory courses, under direct instructional supervision. This course may be repeated for a maximum of 6 credits and may not be used to replace any specific course requirements of the Biology major other than elective units. Statements of planned teaching assistance and tutoring activities required. Pre: consent of the supervising instructor and the department chair.
BIOL x94 Special Topics in Subject Matter (Arr.) (IO) Special topics chosen by the instructor. Course content will vary. May be repeated for credit, provided that a different topic is studied. Additional requirements may apply depending on subject and topic.
BIOL x99 Directed Studies (Arr.) (IO) Statement of planned reading or research required. Pre: instructor’s consent.
- Accounting (ACC) Courses
- Administration of Justice (AJ) Courses
- Agribusiness (AGBU) Courses
- Agricultural Economics (AGEC) Courses
- Agricultural Engineering (AGEN) Courses
- Agriculture (AG) Courses
- Agronomy (AGRN) Courses
- Animal Science (ANSC) Courses
- Anthropology (ANTH) Courses
- Aquaculture (AQUA) Courses
- Art (ART) Courses
- Astronomy (ASTR) Courses
- Biology (BIOL) Courses
- Business (BUS) Courses
- Chemistry (CHEM) Courses
- Chinese (CHNS) Courses
- Civil Engineering (CE) Courses
- Communication (COM) Courses
- Computer Science (CS) Courses
- Dance (DNCE) Courses
- Drama (DRAM) Courses
- Economics (ECON) Courses
- Education (ED) Courses
- Electrical Engineering (EE) Courses
- Engineering (ENGR) Courses
- English (ENG) Courses
- English as a Second Language (ESL) Courses
- Entomology (ENTO) Courses
- Environmental Science/Studies (ENSC) Courses
- Filipino (FIL) Courses
- Finance (FIN) Courses
- Food Science and Technology (FDSC) Courses
- Forestry (FOR) Courses
- French (FR) Courses
- Geography (GEOG) Courses
- Geology (GEOL) Courses
- Hawaiian Language (HAW) Courses
- Hawaiian Studies (HWST) Courses
- History (HIST) Courses
- Honors (HON) Courses
- Horticulture (HORT) Courses
- Interdisciplinary Studies (IS) Courses
- Japanese (JPNS) Courses
- Japanese Studies (JPST) Courses
- Keʻelikōlani Anthropology (KANT) Courses
- Keʻelikōlani Education (KED) Courses
- Keʻelikōlani Hawaiian Language (KHAW) Courses
- Keʻelikōlani Hawaiian Studies (KHWS) Courses
- Keʻelikōlani Indigenous Language (KLAN) Courses
- Keʻelikōlani Indigenous Studies (KIND) Courses
- Kinesiology (KES) Courses
- Language (LANG) Courses
- Latin (LATN) Courses
- Linguistics (LING) Courses
- Management (MGT) Courses
- Marine Science (MARE) Courses
- Marketing (MKT) Courses
- Mathematics (MATH) Courses
- Military Science (MSL) Courses
- Music (MUS) Courses
- Natural Resources (NRES) Courses
- Nursing (NURS) Courses
- Philosophy (PHIL) Courses
- Physics (PHYS) Courses
- Plant Pathology (PPTH) Courses
- Plant Physiology (PPHY) Courses
- Political Science (POLS) Courses
- Psychology (PSY) Courses
- Quantitative Business Analysis (QBA) Courses
- Sociology (SOC) Courses
- Soil Science (SOIL) Courses
- Spanish (SPAN) Courses
- Tourism (TOUR) Courses
- University (UNIV) Courses
- Women’s Studies (WS) Courses